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evolution of fish : ウィキペディア英語版
evolution of fish


The evolution of fish began about 530 million years ago during the Cambrian explosion. Early fish from the fossil record are represented by a group of small, jawless, armoured fish known as ostracoderms. Jawless fish lineages are mostly extinct. An extant clade, the lampreys may approximate ancient pre-jawed fish. The first jaws are found in Placoderm fossils. The diversity of jawed vertebrates may indicate the evolutionary advantage of a jawed mouth. It is unclear if the advantage of a hinged jaw is greater biting force, improved respiration, or a combination of factors. The evolution of fish is not studied as a single event since fish do not represent a monophyletic group but a paraphyletic one (by exclusion of the tetrapods).
==Overview==

Fish may have evolved from an animal similar to a coral-like sea squirt, whose larvae resemble early fish in important ways. The first ancestors of fish may have kept the larval form into adulthood (as some sea squirts do today), although perhaps the reverse is the case. Vertebrates, among them the first fishes, originated about 530 million years ago during the Cambrian explosion, which saw the rise in organism diversity.
The first ancestors of fish, or animals that were probably closely related to fish, were ''Pikaia'', ''Haikouichthys'' and ''Myllokunmingia''. These three genera all appeared around 530 Ma. ''Pikaia'' had a primitive notochord, a structure that could have developed into a vertebral column later. Unlike the other fauna that dominated the Cambrian, these groups had the basic vertebrate body plan: a notochord, rudimentary vertebrae, and a well-defined head and tail. All of these early vertebrates lacked jaws in the common sense and relied on filter feeding close to the seabed.
These were followed by indisputable fossil vertebrates in the form of heavily armoured fishes discovered in rocks from the Ordovician Period 500–430 Ma. The colonisation of new niches resulted in diversification of body plans and sometimes an increase in size. The Devonian Period (395 to 345 Ma) brought in such giants as the placoderm ''Dunkleosteus'', which could grow up to seven meters long, and early air-breathing fish that could remain on land for extended periods. Among this latter group were ancestral amphibians.
The first jawed vertebrates appeared in the late Ordovician and became common in the Devonian, often known as the "Age of Fishes". The two groups of bony fishes, the actinopterygii and sarcopterygii, evolved and became common. The Devonian also saw the demise of virtually all jawless fishes, save for lampreys and hagfish, as well as the Placodermi, a group of armoured fish that dominated much of the late Silurian. The Devonian also saw the rise of the first labyrinthodonts, which was a transitional between fishes and amphibians.
The reptiles appeared from labyrinthodonts in the subsequent Carboniferous period. The anapsid and synapsid reptiles were common during the late Paleozoic, while the diapsids became dominant during the Mesozoic. In the sea, the bony fishes became dominant. The dinosaurs gave rise to the birds in the Jurassic. The demise of the dinosaurs at the end of the Cretaceous promoted expansion of the mammals, which had evolved from the therapsids, a group of synapsid reptiles, during the late Triassic Period.
The later radiations, such as those of fish in the Silurian and Devonian periods, involved fewer taxa, mainly with very similar body plans. The first animals to venture onto dry land were arthropods. Some fish had lungs and strong, bony fins and could crawl onto the land also.

抄文引用元・出典: フリー百科事典『 ウィキペディア(Wikipedia)
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